G. Piovesan et al., TAXONOMIC EVALUATIONS OF THE GENUS PINUS (PINACEAE) BASED ON ELECTROPHORETIC DATA OF SALT SOLUBLE AND INSOLUBLE SEED STORAGE PROTEINS, Plant systematics and evolution, 186(1-2), 1993, pp. 57-68
Electrophoretic studies on seed storage proteins of various gymnosperm
species showed that both salt soluble and insoluble protein fractions
give taxonomic information. Among species of the genus Pinus evident
differences are found between the subgenera Haploxylon (Strobus) and D
iploxylon (Pinus). In sect. Strobus the two subsectt. Strobi and Cembr
ae are readily distinguished from P. bungeana (sect. Parrya) which app
ears to fall in an intermediate position between haploxyl and diploxyl
pines. Among the species of subg. Diploxylon, ''mountain pines'' show
very similar protein patterns in agreement with their recent speciati
on (Quaternary).