Using techniques from supergravity and dimensional reduction, we study
the full isometry algebra of Kahler and quaternionic manifolds with s
pecial geometry. These two varieties are related by the so-called c-ma
p, which may be understood from dimensional reduction of supergravity
theories or by changing chirality assignments in the underlying supers
tring theory. An important subclass, studied in detail, consists of th
e spaces that follow from real special spaces using the so-called r-ma
p. We generally clarify the presence of ''extra'' symmetries emerging
from dimensional reduction and give the conditions for the existence o
f ''hidden'' symmetries. These symmetries play a major role in our ana
lysis. We specify the structure of the homogeneous special manifolds a
s coset spaces G/H. These include all homogeneous quaternionic spaces.