Va. Tucker, GLIDING BIRDS - REDUCTION OF INDUCED DRAG BY WING TIP SLOTS BETWEEN THE PRIMARY FEATHERS, Journal of Experimental Biology, 180, 1993, pp. 285-310
1. The feathers at the wing tips of most birds that soar over land sep
arate both horizontally and vertically in flight to form slotted tips.
The individual feathers in the slotted tips resemble the winglets use
d on the wing tips of some aircraft to reduce induced drag. 2. A wing
that produces lift leaves a pair of vortex sheets in its wake. Wing th
eory shows that winglets can reduce the kinetic energy left in the vor
tex sheets, and hence the induced drag, by spreading vorticity both ho
rizontally and vertically. 3. This paper describes the aerodynamic for
ces on a wing made of a base wing and different wing tips. The feather
ed wing tip was slotted and was made of four primary feathers from a H
arris' hawk (Parabuteo unicinctus). The Clark Y tip was unslotted and
was made of balsa wood shaped to a Clark Y aerofoil. The balsa feather
tip was slotted and was made of three balsa wood wings shaped like fe
athers. 4. The base wing in a wind tunnel at an air speed of 12.6 m s-
1 generated upwash angles as high as 15-degrees at the end of the wing
when the angle of attack of the wing was 10.5-degrees. The feathered
tip responded to upwash by increasing its lift to drag ratio (L/D) by
107%, from 4.9 to 10.1, as the angle of attack of the base wing increa
sed from 4-degrees to 14-degrees. The L/D values of the balsa feather
tip and the Clark Y tip increased by 49% and 5%, respectively, for the
same change in angle of attack. 5. With the angle of attack of the ba
se wing fixed at 13-degrees, changing the angle of attack of the wing
tip changed the drag of the base wing. The drag of the base wing incre
ased by 25% as the angle of attack of the Clark Y tip increased from 0
-degrees to 15-degrees. The base wing drag decreased by 6% for the sam
e change in the angle of attack of the feathered tip. 6. The total dra
g of the wing with the feathered tip was 12% less than that of a hypot
hetical wing with the same lift and span, but with tip feathers that d
id not respond to upwash at the end of the base wing. This value is co
nsistent with wing theory predictions on drag reduction from winglets.
7. Wings with the tip and the base wing locked together had lift and
drag that increased with increasing base wing angle of attack, as expe
cted for conventional wings. Span factors were calculated from these d
ata - a large span factor indicates that a wing has low induced drag f
or a given lift and wing span. The wing with the Clark Y tip had a spa
n factor that decreased from 1 to 0.75 as the angle of attack of the b
ase wing increased. Over the same range of angle of attack. the span f
actor of the wing with the feathered tip remained constant at 0.87. As
the angle of attack of this wing increased, aerodynamic forces spread
the feathers vertically to form slots. With fully formed slots, the w
ing had a higher span factor than the wing with the unslotted Clark Y
tip. 8. Flow visualization with helium-filled bubbles showed that the
addition of two winglets to the tip of a model wing spread vorticity b
oth horizontally and vertically in the wake of the tip. 9. These obser
vations taken together provide strong evidence that the tip slots of s
oaring birds reduce induced drag in the sense that the separated tip f
eathers act as winglets and increase the span factor of the wings.