The evolution of aggregation in seabirds is usually attributed to pred
ation pressure, although many authors have argued for the role of fora
ging constraints to be considered. Discriminating between factors that
result in group living and characteristics arising from group living
has been regarded as an insoluble problem; yet it is possible to test
the predictions of the different models proposed to explain colonial n
esting. The resulting explanation suggests that seabird aggregations h
ave evolved in order to utilize a dispersed and unpredictable food sup
ply. As such, colonial nesting leads to increased vulnerability to pre
dation, rather than being a strategy adopted to combat it.