A cross-fostering experiment demonstrated that tarsus length of Europe
an starlings (Sturnus vulgaris) was heritable. The tarsus length of th
e foster-parent had no effect. A full-sib resemblance analysis showed
that sibs were much more similar in tarsus length than explained by he
ritability alone. This was partly due to an effect of female mating st
atus on offspring tarsus length. When nestling growth was retarded in
secondary females' nests due to reduced male assistance, the increased
environmental variation in tarsus length masked the heritability.