Watts (1969) described the population cycle of the Wood Mouse Apodemus
sylvaticus as a decline in numbers in spring, a stable period in summ
er and an increase phase in autumn. Aggression by adult males was cons
idered an important aspect of population regulation limiting male surv
ival in spring and juvenile recruitment in summer. However, recent stu
dies of range behaviour and experimental investigations suggest that m
ale A. sylvaticus are not relevant in population regulation and that t
he principal regulating factors act on female reproductive success. Fe
male reproductive activity is determined by spatial interactions which
, in turn, are determined primarily by the density of females and the
proportion of female immigrants in the population. The only impact adu
lt males may have on population size is through infanticide and compet
ition with the females for food.