Birds are highly mobile animals and we need to understand the factors
and processes that determine numbers throughout their range and annual
life cycle. Most avian studies have concentrated on birds using one l
ocality at a particular time of year, but the numbers breeding or wint
ering in any one locality may be affected by the size of the greater p
opulation of which the local group forms a part, as well as by local c
onditions. Equilibrium theory provides a useful framework within which
to analyse populations at this larger scale, although the meaning and
usefulness of some of the concepts used for investigating local group
s may change. New approaches are required in which mathematical models
are used to integrate the results of studies on the behavioural and p
opulation ecology of local groups with those of large-scale monitoring
programmes of the kind conducted by the BTO.