The lengths of food chains within ecosystems have been thought to be l
imited either by the productivity of the ecosystem or by the resilienc
e of that ecosystem after perturbation. Models based on ecological ene
rgetics that follow the form of Lotka-Volterra equations and equations
that include material (detritus) recycling show that productivity and
resilience are inextricably interrelated. The models were initialized
with data from 5- to 10-year studies of actual soil food webs. Estima
tes indicate that most ecological production worldwide is from ecosyst
ems that are themselves sufficiently productive to recover from minor
perturbations.