New Zealand Rifleman Acanthisitta chloris females are larger and brown
er than males (which are green dorsally), have a relatively larger hin
d claw and a longer, more decurved bill. Using three criteria for dist
inguishing sexual selection from ecological influences on sexual dimor
phism, we suggest that sexual dimorphism in the Rifleman is maintained
by ecological factors. When energy requirements were high (feeding fl
edglings), female and male Rifleman foraged separately. When energy re
quirements were low (non-breeding), females and males fed together. Pl
umage differences between the sexes were correlated with the different
habitats in which Riflemen foraged when energy demands were high.