C. Vandenberg et Cp. Ellington, THE VORTEX WAKE OF A HOVERING MODEL HAWKMOTH, Philosophical transactions-Royal Society of London. Biological sciences, 352(1351), 1997, pp. 317-328
Visualization experiments with Manduca sexta have revealed the presenc
e of a leading-edge vortex and a highly three-dimensional flow pattern
. To further investigate this important discovery, a scaled-up robotic
insect was built (the 'flapper') which could mimic the complex moveme
nts of the wings of a hovering hawkmoth. Smoke released from the leadi
ng edge of the flapper wing revealed a small but strong leading-edge v
ortex on the downstroke. This vortex had a high axial flow velocity an
d was stable, separating from the wing at approximately 75% of the win
g length. It connected to a large, tangled tip vortex, extending back
to a combined stopping and starting vortex from pronation. At the end
of the downstroke, the wake could be approximated as one vortex ring p
er wing. Based on the size and velocity of the vortex rings, the mean
lift force during the downstroke was estimated to be about 1.5 times t
he body weight of a hawkmoth, confirming that the downstroke is the ma
in provider of lift force.