Main et al. (1996) synthesized evidence that supported or refuted three hyp
otheses that have emerged as the most common explanations for sexual segreg
ation in ungulates. They provided new insights by evaluating a broad range
of studies, and they identified promising topics for future research. Howev
er, Main et al.'s (1996) analyses may lead to further confusion about impor
tant links between foraging by ungulates and mechanisms that can account fo
r sexual segregation. To clarify those links, I summarize information on fo
raging processes central to understanding and evaluating hypotheses on diet
selection, habitat use, and sexual segregation by ungulates. I reevaluate
evidence presented by Main et al. (1996) and identify key differences in fe
eding by browsers and grazers that affect the interpretation of morphologic
al characteristics. To improve future studies of sexual segregation, I prov
ide additional suggestions that incorporate recent findings on feeding by u
ngulates.