The amazing diversity of extant photosynthetic eukaryotes is largely a resu
lt of the presence of formerly free-living photosynthesizing organisms that
have been sequestered by eukaryotic hosts and established as plastids in a
process known as endosymbiosis. The evolutionary history of these endosymb
iotic events was traditionally investigated by studying ultrastructural fea
tures and pigment characteristics but in recent years has been approached u
sing molecular sequence data and gene trees. Two important developments, mo
re detailed studies of members of the Cyanobacteria (from which plastids ul
timately derive) and the availability of complete plastid genome sequences
from a wide variety of plant and algal lineages, have allowed a more accura
te reconstruction of plastid evolution.