Six sympatric species of 5-million-year-old (Late Hemphillian) horses from
Florida existed during a time of major global change and extinction in terr
estrial ecosystems. Traditionally, these horses were interpreted to have fe
d on abrasive grasses because of their high-crowned teeth. However, carbon
isotopic and tooth microwear data indicate that these horses were not ail C
-4 grazers but also included mixed feeders and C-3 browsers. The late Hemph
illian Florida sister species of the modern genus Equus was principally a b
rowser, unlike the grazing diet of modern equids. Late Hemphillian horse ex
tinctions in Florida involved two grazing and one browsing species.