Adequate forage amounts in both physical and chemical forms are necessary f
or proper ruminal function in dairy cows. Under conditions in which total a
mounts of forage or particle size of the forage are reduced, cows spend les
s time ruminating and have a decreased amount of buoyant digesta in the rum
en. These factors reduce saliva production and allow ruminal pH to fall, de
pressing activity of cellulolytic bacteria and causing a prolonged period o
f low ruminal pH. Insufficient particle size of the diet decreases the rumi
nal acetate-to-propionate ratio and reduces ruminal pH. The mean particle s
ize of the diet, the variation in particle size, and the amount of chemical
fiber (i.e., NDF or ADF) are all nutritionally important for dairy cows. D
efining amounts and physical characteristics of fiber is important in balan
cing dairy cattle diets. Because particle size plays such an important role
in digestion and animal performance, it must be an important consideration
from harvest through feeding. Forages should not be reduced in particle si
ze beyond what is necessary to achieve minimal storage losses and what can
be accommodated by existing equipment. Forage and total mixed ration (TMR)
particle sizes are potentially reduced in size by all phases of harvesting,
storing, taking out of storage, mixing, and delivery of feed to the dairy
cow. Mixing feed causes a reduction in size of all feed particles and is di
rectly related to TMR mixing time; field studies show that the longest part
icles (> 27 mm) may be reduced in size by 50%. Forage and TMR particle size
as fed to the cows should be periodically monitored to maintain adequate n
utrition for the dairy cow.