The concept of preadaptation, though useful, continues to trouble evolution
ary scientists. Usually, it is treated as if it were really adaptation, pro
mpting such diverse theorists as Gould and Vrba, and Dennett to suggest its
removal from evolutionary theory altogether. In this paper, I argue that t
he as-if sense is ill-founded, and that the sense of preadaptation as a pro
cess may be defended as unequivocal and generally useful in evolutionary ex
planations, even in such problem areas as human evolution.