Ld. Vanvleck, CALCULATION OF BREED DIRECT AND MATERNAL GENETIC FRACTIONS AND BREED SPECIFIC DIRECT AND MATERNAL HETEROZYGOSITY FOR CROSSBREEDING DATA, BRAZ J GENE, 20(4), 1997, pp. 639-644
Teaching, research, and herd breeding applications may require calcula
tion of breed additive contributions for direct and maternal genetic e
ffects and fractions of heterozygosity associated with breed specific
direct and maternal heterosis effects. These coefficients can be obtai
ned from the first NE rows of a pseudo numerator relationship matrix w
here the first NE rows represent fractional contributions by breed to
each animal or group representing a specific breed cross. The table be
gins with an NE x NE identity matrix representing pure breeds. Initial
animals or representative crosses must be purebreds or two-breed cros
ses. Parents of initial purebreds are represented by the corresponding
column and initial two-breed cross progeny by the two corresponding c
olumns of the identity matrix. Af ter that, usual rules are used to ca
lculate the NE column entries corresponding to breeds for each animal.
The NE entries are fractions of genes expected to be contributed by e
ach of the pure breeds and correspond to the breed additive direct fra
ctions. Entries in the column corresponding to the dam represent breed
additive maternal fractions. Breed specific direct heterozygosity coe
fficients are entries of an NE x NE matrix formed by the outer product
of the two NE by 1 columns associated with sire and dam of the animal
. One minus sum of the diagonals represents total direct heterozygosit
y. Similarly, the NE x NE matrix formed by the outer product of column
s associated with sire of dam and dam of dam contains breed specific m
aternal heterozygosity coefficients. These steps can be programmed to
create covariates to merge with data. If X represents these coefficien
ts for all unique breed crosses, then the reduced row echelon form fun
ction of MATLAB or SAS can be used on X to determine estimable functio
ns of additive breed direct and maternal effects and breed specific di
rect and maternal heterosis effects.