Numerous mutualistic associations between phototrophs and fungi exist
in the extant land biota. Some are widespread, such as lichens and myc
orrhizae, but some are less well known or restricted to special ecolog
ical conditions, such as endophytes in plants and algae. Recent molecu
lar data and fossils suggest that associations arose repeatedly and th
at some of them are ancient, and even ancestral in the case of land pl
ants. Mutualism, that provides various adaptations to terrestrial cons
traints, may have played a crucial role during terrestrialization and
evolution of land phototrophs.