Molecular and genetic studies show that the underlying mechanisms cont
rolling flower development are largely conserved in distantly related
dicotyledonous plant species. These studies have identified early-acti
ng genes that promote the formation of floral meristems and later-acti
ng genes that determine the fate of floral organ primordia. The floral
meristem identity genes lie at the top of a regulatory hierarchy that
leads to the activation of the organ identity genes. Subsequent regul
atory interactions between the organ identity genes restrict their dom
ains of activity. The resulting spatial domains of gene activity allow
cells within the floral meristem to assess their position and, hence,
to differentiate into the appropriate organ type.