Plagiolaria nandanensis is a zonal index-fossil from the Lower Devonia
n of Nandan, Guangxi, China. Strata of this age yield numerous exuvies
of trilobites. These exuvies show that the break of the dorsal exoske
leton in P. nandanensis started with the separation of the thorax and
the cephalon; the animal shed the hypostoma and crawled forward, lifti
ng up the cephalon and then turning it upside down. Sometimes, during
a final stage, the thorax and the pygidium cleaved apart. Moulting in
Plagiolaria nandanensis represents the mode of moulting in small-eyed
Phacopids.