J. Rohacek, TAXONOMIC LIMITS, PHYLOGENY AND HIGHER CLASSIFICATION OF ANTHOMYZIDAE(DIPTERA), WITH SPECIAL REGARD TO FOSSIL RECORD, European journal of entomology, 95(1), 1998, pp. 141-177
Taxonomic limits of the family Anthomyzidae are prescribed. Two fossil
genera are affirmed, viz. Protanthomyza Hennig, 1965 (Baltic amber) a
nd Grimalantha gen. n. (type species: G. vulnerata sp. n.) described f
rom Dominican amber. Fourteen extant genera are recognized, including
Chamaebosca Speiser, 1903 (= Penquistus Kieffer, 1906 syn. n.) and Apt
erosepsis Richards, 1962. New diagnoses of the latter two genera and r
edescriptions of their type species are given and their relationships
are discussed. Chamaebosca cursor (Kieffer, 1906) becomes a new combin
ation. The monotypic genus Echidnocephalodes lodes Sabrosky, 1980 is r
emoved from Anthomyzidae, newly diagnosed and its type species E. barb
atus (Lamb, 1914) redescribed and a lectotype designated. Echidnocepha
lodes is considered to be related to Periscelididae and/or Aulacigastr
idae, particularly to those genera with symmetrical male postabdomen.
The inferred phylogeny of the Anthomyzidae, based on cladistic analysi
s, is presented. The Opomyzidae are confirmed as a sister-group of the
Anthomyzidae, while Protanthomyza is found to be the most primi tive
anthomyzid genus forming a sister-group to all recent genera plus the
fossil Grimalantha gen. n. The monophylies of the latter group of gene
ra, and of the Anthomyzidae as a whole, are demonstrated. The genus Pr
otanthomyza is classified in a new subfamily Protanthomyzinae, and all
remaining genera are placed in the subfamily Anthomyzinae Frey, 1921.
An annotated world checklist of the family Anthomyzidae is appended.