The marine bivalved Brachiopoda are abundant throughout the geological
record and have apatitic (CaPO4-rich) or calcitic (CaCO3-rich) shells
. Vesicles covering the larval valves of living apatitic-shelled disci
nids contain tablets of silica. The tablets are cemented into close-pa
cked mosaics by spherular apatite in glycosaminoglycans. They are usua
lly lost as vesicles degrade but leave imprints on the underlying apat
itic shell. Similar imprints ornament larval surfaces of some of the e
arliest Paleozoic apatitic-shelled brachiopods and may also be indicat
ors of siliceous biomineralization.