Plant communities of trampled soil dominated by plants characterized b
y the C4-assimilation syndrome were investigated in Europe. These spec
ies, belonging to genera such as Chamaesyce, Amaranthus, Eleusine, Era
grostis and Setaria, are thermophilous, late-germinating, prostrate he
rbs or grasses. The centre of their distribution is in the (Sub)Tropic
s. A syntaxonomic revision of the phytosociological material from Euro
pe (incl. the Macaronesian Archipelago) revealed three alliances: the
Euphorbion prostratae from Spain, the Polycarpo-Eleusinion indicae fro
m Italy, and Slovenian and Croatian Istria, and the Eragrostio-Polygon
ion arenastri from temperate regions of Europe. The latter two syntaxa
are described as new. All three alliances belong to the order Eragros
tietalia (class Stellarietea mediae). Vicarious (ecologically analogou
s) communities occur also in southern Africa, eastern Asia and North A
merica. The communities studied in the present paper are considered to
be an impoverished form of highly diversified trampled plant communit
ies typical of (sub)tropical areas.