A CLADISTIC-ANALYSIS OF THE TRIBE EPISCIEAE (GESNERIACEAE) BASED ON NDHF SEQUENCES - ORIGIN OF MORPHOLOGICAL CHARACTERS

The neotropical members of Gesneriaceae are characterized by numerous characters that appear to be adaptations to life in the wet tropics. A mong these are epiphytism, anisophylly, hummingbird-pollination, and b ird-dispersal of seeds. Many of these characters have diversified to a broad extent in the single tribe Episcieae. This analysis investigate s the phylogenetic relationships among genera of Episcieae and also ex amines the evolutionary origin of several characters found within this tribe such as fruit characters, epiphytism, chromosome numbers, and t ubers. Ail genera of the tribe were included in the analysis except th e Guyana endemic Rhoogeton and the Central American Oerstedina. Larger genera such as Columnea and Episcia were represented by several speci es with attempts to include members of the different sections of these genera. Columnea appears to be a monophyletic group but its relations hip to Drymonia and Alloplectus is not resolved. The latter genera may need to be included in Columnea to make it strictly monophyletic. Neo mortonia is polyphyletic with one species in the Columnea dade and ano ther with Episcia. The anomalous position of some Paradrymonia indicat e a more thorough analysis of this genus is necessary to resolve phylo genetic relationships.