The annual cycle of adult butterflies was studied at seven rocky outcr
ops located along an altitudinal gradient in a northern Spanish mounta
in area (Picos de Europa) with a Eurosiberian climate in 1993 and 1995
. Both total abundance and number of species of butterflies peaked in
summer at all study sites. The cumulative species number uniformly inc
reased over the season until late summer. These results imply that but
terflies concentrated their activity when temperatures were higher in
the year. Regardless of their diversity and total abundance, timing of
the flight period was later for assemblages at high altitude than for
assemblages at low altitude. A similar trend in increasing delay in f
light period was found when data were analysed separately for each spe
cies. The duration of the flight period of assemblages was correlated
with abundance in 1993, but it was correlated with altitude in 1995. M
ost species tended to show shorter flight periods at high altitudes in
both study years. These altitudinal changes in timing of adult flight
are likely to be linked with effects of temperature on developmental
rates of early stages of the life cycle. Changes in duration of the fl
ight period, however, might be partly related to variability in specie
s abundance.