A hammer mill removed most of the lemmas, paleas, and pericarps from c
heat florets. Typically, the cuticular layer of the testa was the only
remaining intact layer, and damage to the embryos and endosperm was s
evere. A roller mill disrupted tissue organization of lemmas, paleas,
and outer layers of the caryopses primarily at the cuts. Large gaps be
tween the aleurone layer and testa, between testa and pericarp, and be
tween the scutellum and endosperm were created. In the field, germinat
ion of mill-damaged florets was reduced, and florets exhibited progres
sive degradation the longer they were buried. Nematodes and fungi pene
trated the cuticular layer of mill-damaged seed. Attaching a hammer mi
ll or a roller mill to a grain combine to treat cheat seed before it i
s returned to the field could provide a novel method of cheat control.