F. Dupuis et Rp. Dechambre, SYSTEMATIC AND PHYLOGENY OF THE GENUS ACT INOBOLUS WESTWOOD, 1841 (COLEOPTERA, SCARABAEOIDEA, DYNASTIDAE), Annales de la Societe entomologique de France, 34(1), 1998, pp. 33-43
The genus Actinobolus Westwood is revised : 3 species (A. alvarengai,
sp. n., A. pilosus, sp. n., A. spiniformis, sp. n.) are described;A. t
alpipes Ohaus, 1915 is synonymized with A. radians Westwood 1841; an i
dentification key, revised diagrams of systematic characteristics and
distribution maps are provided. A phylogenetic study, based an cladist
ics, enables to discuss the main characteristics of the 8 species and
to present a phylogenetic essay of the genus. A review of the main bio
logical data establishes the association of the group with termitories
.