Vertebrate fossils and continental sediments provide a rich record of
variations in the isotopic composition of surface environments. To int
erpret these records, a greater understanding of isotopic sources, as
well as fractionations associated with animal physiology, soil geochem
istry, and diagenesis, has been essential. Tooth enamel and fish otoli
ths yield subannual records of surface environments, whereas soil mine
rals may integrate signals over many thousands of years. Carbon isotop
e variations in fossil vertebrates and soils record changes in the str
ucture of vegetation and the isotope composition and concentration of
atmospheric CO2. Oxygen isotope variations may be indirectly related t
o climate, through reconstruction of the oxygen isotope composition of
meteoric water, or directly related to temperature, through applicati
on of oxygen isotope paleothermometry to soil minerals or otoliths. In
Africa, nitrogen isotope variations show promise as a proxy for rainf
all abundance, though the generality of this association elsewhere has
not been demonstrated.