COLLAPSE OF ISERTIEAE, REESTABLISHMENT OF MUSSAENDEAE, AND A NEW GENUS OF SABICEEAE (RUBIACEAE) - PHYLOGENETIC-RELATIONSHIPS BASED ON RBCL DATA

The circumscription of the Isertieae has been under debate for a long time and recently a phylogeny based on morphological data has been pre sented (ANDERSSON 1996), contradicting the classification of ROBBRECHT (1988, 1993). Our investigation of molecular data neither supports th e phylogeny of ANDERSSON nor the classification of ROBBRECHT, but inst ead indicates totally new relationships of Isertieae, Mussaendeae, and Sabiceeae. The Isertieae are a bigeneric tribe of subfam. Cinchonoide ae, while Mussaendeae and Sabiceeae are two separate tribes of subfam. Ixoroideae. We have also referred a species from Socotra (Yemen) with disputed position to the tribe Sabiceeae and we place it in a new gen us, Tamridaea, with the single species T. capsulifera comb. nov. New r bcL sequences of 20 taxa are presented and analysed, from Gentianaceae : Gentianella; from Loganiaceae: Spigelia; and from Rubiaceae: Amphida sya, Aoranthe, Chomelia, Coussarea, Gonzalagunia, Heinsia, Hippotis, I sertia (three taxa), Mussaenda, Pseudomussaenda, Pseudosabicea, Rondel etia, Sabicea, Schraclera, Tamridaea, and Virectaria.