The evolutionary history of the living hominoids has remained elusive
despite years of exploration and the discovery of numerous Miocene fos
sil ape species. Part of the difficulty can be attributed to the chang
ing nature of our views about the course of hominoid evolution. In the
1950s and 1960s, individual Miocene taxa were commonly viewed as the
direct ancestors of specific living ape species, suggesting an early d
ivergence of the modern lineages.(1-5) However, in most cases, the Mio
cene forms were essentially ''dental apes,'' resembling extant species
in dental and a few cranial features, but possessing more primitive p
ostcranial features that suggested arboreal quadrupedalism rather than
suspensory habits. With the introduction of molecular methods of phyl
ogenetic reconstruction and the increasing use of cladistic analysis,
it has become apparent that the radiation leading to the modern homino
ids was somewhat more recent than had been believed, and that most of
the Miocene hominoid species had little to do with the evolutionary hi
story of the living apes.(6)