Counts of taxonomic diversity are the prevailing standards for documen
ting Large-scall patterns of evolution in the fossil record, However,
the secular pattern of relative ecological importance between the bryo
zoan clades Cyclostomata and Cheilostomata is not reflected fully in c
ompilations of generic diversity or within-fauna species richness, and
the delayed ecological recovery of the Cheilostomata after the mass e
xtinction at the Cretaceous-Tertiary boundary is missed entirely. Thes
e observations demonstrate that evolutionary success and ecological do
minance can be decoupled and profoundly different, even over tons of m
illions of years.