NEW FAN WORM GENERA AND SPECIES (POLYCHAETA, SABELLIDAE, FABRICIINAE)FROM THE WESTERN PACIFIC, AND CLADISTIC RELATIONSHIPS AMONG GENERA

Five new species are described from among the genera Fabriciola Friedr ich. 1939 tone species from Okinawa), Novafabricia Fitzhugh, 1990 (two species from Australia and Papua New Guinea, respectively), and two n ew monotypic genera, Brifacia gen. n. (Australia), and Pseudoaugenerie lla gen. n. (Okinawa). All species are known only from the intertidal. Fabriciola rubra sp, n, is a member of the species complex with red p ersitomial and pygidial eyes, inferior thoracic flagellate setae, and abdominal pin-head setae. Novafabricia lab,us sp. n., and N. exiguus s p. n., are distinguished from other species based on the distribution of inferior thoracic pseudospatulate setae. Brifacia metastellaris sp. n., resembles Fabricia stellaris (Muller, 1774) with regard to anteri or peristomial ring collar construction and distribution of pseudospat ulate setae, but differs in having abdominal uncini with relatively sh ort manubria. Pseudoaugeneriella uniranta sp. n., most closely resembl es species of Augeneriella Banse, 1956 in general form. but has unbran ched vascularized ventral filamentous appendages. Revised keys to fabr iciin genera and selected species are provided. Cladistic relationship s of these new genera and species to other Fabriciinae are examined an d the status of some characters are reviewed. Whereas previous analyse s have supported a monophyletic Augeneriella on the basis of branched, vascularized ventral filamentous appendages, present results show Aug eneriella paraphyletic relative to Fabricia Blainville, 1828, Novafabr icia, Fabricinuda Fitzhugh, 1990 Parafabricia Fitzhugh, 1992 and Brifa cia. Similarly, some topologies show Novafabricia, which has only been defined by the presence of poorly developed dorsal lips, to be paraph yletic relative to Fabricia, Fabricinuda, Parafabricia, and Brifacia. The cladistic analyses dal however, support recognition of Bn;facia an d Pseudoaugeneriella. With the addition of Brifacia, the presence of u nbranched, vascularized appendages is plesiomorphic for the Fabriciina e, which has not been previously hypothesized. (C) 1998 The Norwegian Academy of Science and Letters.