M. Ruta et C. Bartels, A REDESCRIPTION OF THE ANOMALOCYSTITID MITRATE RHENOCYSTIS-LATIPEDUNCULATA FROM THE LOWER DEVONIAN OF GERMANY, Palaeontology, 41, 1998, pp. 771-806
The anomalocystitid mitrate Rhenocystis latipedunculata, from the Lowe
r Devonian Hunsruckschiefer of Rhineland, Germany, is reconstructed an
d redescribed. Rhenocystis is characterized by transverse, terrace-lik
e ridges on two antero-posteriorly elongate, postero-lateral areas of
the dorsal head skeleton and on the posterior third of the ventral hea
d skeleton; the relatively small size of the ventral plates of the sec
ond transverse row; a suture between the mid-ventral plates of the fir
st and third row; the relatively large size of the placocystid plate;
the presence of rocking articulations between dorsal and ventral fore
tail plates; a transversely expanded and recumbent anterior styloid bl
ade; a robust, spike-like posterior blade; and four morphologically di
stinct regions in the hind tail. Rhenocystis closely resembles Placocy
stites forbesianus from the middle Silurian of England and Victoriacys
tis wilkinsi from the upper Silurian of Australia, with which if forms
a clade within the anomalocystitids of boreal type.