O. Rieppel, THE SYSTEMATIC STATUS OF HANOSAURUS-HUPEHENSIS (REPTILIA, SAUROPTERYGIA) FROM THE TRIASSIC OF CHINA, Journal of vertebrate paleontology, 18(3), 1998, pp. 545-557
Hanosaurus hupehensis was believed to represent a Middle Triassic thal
attosaur from the People's Republic of China. Redescription of the hol
otype and only known specimen, and cladistic analysis of its interrela
tionships, shows Hanosaurus to represent a distinct pachypleurosaur. T
he two Chinese pachypleurosaurs, Keichousaurus and Hanosaurus, form tw
o consecutive sister-groups to European pachypleurosaurs, strengthenin
g the Asiatic affinities of the clade. Addition of Hanosaurus furtherm
ore results in a modification of pachypleurosaur interrelationships as
previously reconstructed: the pachypleurosaurs from the Germanic Tria
ssic (Anarosaurus, Dactylosaurus) form a monophyletic clade, which is
sister-group to the monophyletic clade of Alpine pachypleurosaurs (Ser
pianosaurus, Neusticosaurus). Thalattosauria are the sister-group of S
auropterygia, the two clades possibly representing a single invasion o
f the Mesozoic sea. However, more detailed knowledge of thalattosaur a
natomy is required to further test the phylogenetic relationships of t
halattosaurs.