This study examines the on-line processing of morphological cues to sentenc
e interpretation in Russian and German with the goal of evaluating the rela
tive impacts of cue availability and cue reliability. Both Russian and Germ
an use the cues of word order, animacy, case-marking, and subject-verb agre
ement to identify the agent of active transitive sentences. However, the av
ailability of the case-marking cue is higher in Russian than in German. Usi
ng a picture-choice paradigm, we contrasted case-marking and animacy in Rus
sian and German. The reaction times showed larger effects of case-marking i
n Russian than in German and effects of animacy in German, but not in Russi
an. These results suggest that the higher the availability of a cue, the la
rger the processing benefits associated with the presence of this cue and t
he smaller the impact of other converging information. A recurrent cascaded
backpropagation network was designed to simulate these effects. The networ
k succeeded in capturing the essential language differences in the reaction
times, thereby illustrating how the statistical properties of cues in a la
nguage can affect the time-course of activation of alternative interpretati
ons during sentence processing.