Cordyceps variabilis has long been erroneously reported to be a pathog
en of beetle larvae; studies of newly collected and herbarium specimen
s revealed that C. variabilis attacks fly larvae, particularly those o
f the family Xylophagidae (Diptera). An emended description of C. vari
abilis is provided and a lectotype designated. The anamorph of C. vari
abilis is reported here for the first time, and is attributed to the h
yphomycetous genus Syngliocladium. An allied species, S. tetanopsis sp
. nov. is reported as a pathogen of larvae of Tetanops myopaeformis, t
he sugar beet root maggot (Diptera: Otitidae). The taxonomy of the gen
us Syngliocladium is briefly reviewed, and the generic concept emended
.