R. Zainol et al., ANATOMICAL ANALYSIS OF DOUBLE-FLOWER MORPHOGENESIS IN A NICOTIANA-ALATA MUTANT, Journal of the American Society for Horticultural Science, 123(6), 1998, pp. 967-972
Anatomical analysis was performed using a double-flowered mutant of Ni
cotiana alata Link & Otto. Flower doubleness resulted from petaloid mo
dification of the androecium. Vascularized petal-like outgrowths arose
from the anther, connective, and filament of the stamen. The vasculat
ure in petaloid outgrowths from the anther and upper part of the filam
ent originated from and was continuous with the vascular bundle of the
filament. In contrast, the vascular bundles formed in the outgrowths
from the lower part of the filament developed independently of the vas
cular bundle of the filament and were not connected to it at any time.
Emergences consisting of epidermal and ground parenchyma tissue and l
acking vascularization arose from the filament.