The recent application of immunohistochemistry and molecular techniques has
revealed that endosperm development depends on a genetic program that comb
ines an ancient process for cellularization (similar to that seen in late P
aleozoic seed ferns) with a mechanism for specifying asymmetric cell fates
that has parallels to signaling processes in mammals. Progress has been fur
ther accelerated by the recent realization that the conserved nature of nuc
lear endosperm development extends beyond the grass species, to include dic
ots, such as Arabidopsis. It is anticipated that these ongoing studies will
provide invaluable tools for the improvement of yield and grain quality in
cereal crops.