K. Marhold, Taxonomic evaluation of the tetraploid populations of Cardamine amara (Brassicaceae) from the Eastern Alps and adjacent areas, BOT HELV, 109(1), 1999, pp. 67-84
Tetraploid populations of C. amara occupying the area of the Eastern Alps a
nd neighbouring areas are described as a new subspecies, C. amara subsp. au
striaca Marhold. The distribution area of these populations largely coincid
es with the area heavily affected by Pleistocene glaciation and it is sugge
sted that the polyploidization event or events occurred during interglacial
or early postglacial periods. Morphometric analysis of diploid and tetrapl
oid populations of C. amara revealed that the new subspecies is morphologic
ally close to diploid C. amara subsp. amara, but it can be separated from t
his taxon by the combination of several morphological characters from among
of which the size of pollen grains is the most important.