The nature of synmetamorphic fluids and their flow is examined in the granu
litic lower crust of Madagascar, part of a Precambrian crustal-scale networ
k of vertical ductile shear zones. Based on three independent data sets - f
ield and satellite mapping, C-, O- and H-isotope geochemistry and gravimetr
y - this crust is divided into three zones: outside of shear zones, minor s
hear zones (<140 km long and 7 km wide), and major shear zones (>350 km lon
g and 20-35 km wide). The major shear zones are rooted in and are controlle
d by the mantle. They tapped mantle-derived CO2 with carbon fluxes of the s
ame order of magnitude as oceanic ridge degassing. One major shear zone sho
ws abundant phlogopite-diopside-apatite-calcite mineralizations (a well kno
wn paragenesis in mantle metasomatism) due to mantle-fluid infiltration and
their interaction with the crust. Carbonatitic magmas possibly collected i
n the major shear zones at the base of the crust and may be the source for
CO2 upwellings as well as other metasomatic agents. Small-scale minor shear
zones were controlled by crustal deformation processes and focused crustal
ly-derived H2O-rich fluids. Pervasive fluid circulation was restricted to t
he vicinity (< 100 m) of synmetamorphic plutons. Fluid absent conditions do
minate everywhere else. Mantle-CO2 flushing is not required for granulite g
enesis but is a consequence of the high associated heat flux. Fluid transfe
r at the mantle/crust interface is controlled by the tectonic setting and t
he associated geothermal gradient. The C- and O- isotope systematics of met
amorphosed carbonates sampled on a regional scale within a known petrologic
al and structural framework are shown to be of great help to identify the d
istribution of major fluid-rock interaction processes associated with plate
tectonics.