The fine structure of the tubulobulbar complex (TBC) of the ram semini
ferous epithelium was studied in immersion-fixed testicular samples ob
tained in autumn (sheep's mating period). The TBC was visible during t
he last two stages of the seminiferous epithelium cycle. It originated
as a tubular invagination of the Sertoli cytoplasm harbouring a compl
ementary evagination from the spermatid head. Later, it expanded into
a bulbous structure, finally becoming detached from the spermatid and
apparently phagocytosed by the Sertoli cell. The significance of this
transient structure is discussed and compared with previous reports fo
r other eutherian mammals.