Mf. Proe et al., Use of vector analysis to determine the effects of harvest residues on early growth of second-rotation Sitka spruce, FOREST ECOL, 122(1-2), 1999, pp. 87-105
Vector analyses have been used to examine the effects of harvest residues o
n the growth of Sitka spruce planted 18 months after clearfelling a first r
otation of the same species. Four treatments were imposed at the time of pl
anting: retention of all residues; retention of fine twigs and foliage; pla
cement of coarse branch material onto plots; and removal of all harvest res
idues. Sample trees were harvested each year for five years after planting
and the biomass and nutrient contents of whole crowns were quantified.
Harvest residues had a large effect on mean tree height (p < 0.001). Coarse
and fine components produced similar increases when compared to plots with
no residues although the response to fine material occurred earlier. After
five years, residues had increased mean tree height by 40%. Residues also
increased the mean dry weight per tree although the response to coarse mate
rial was not significant until the age of five years, at which time trees i
n residues had 166% more biomass (p< 0.001). Tree growth was greatest where
both forms of residue were retained on site.
Interpretation of vector nomograms, foliage concentrations and concentratio
n ratios indicated that the growth response was unlikely to he related to c
hanges in nutrient supply for the first two years of treatment. Early treat
ment effects may have been mediated through changes in weed competition or
microclimate. After three years, treatment effects were likely to have been
related to altered nutrient supply, particularly those for nitrogen and po
tassium. According to vector analyses, potassium was not limiting tree grow
th in whole-tree harvested plots although foliage levels were below critica
l levels for young Sitka spruce and potassium : nitrogen ratios indicated a
nutrient imbalance in these plots. Such an imbalance may arise from the ra
pid loss of potassium from decomposing harvest residues accompanied by a mo
re gradual release of nitrogen that may relate more closely to the nutrient
demand of newly planted trees.
The contention that vector analyses allow a standardized comparison between
treatments without confounding with other factors has to be questioned. In
the present study, different interpretations resulted from analyses based
on comparing samples of different age-classes or from different canopy posi
tions. The method has also been shown to be very sensitive to the choice of
critical test statistic used in assessing each vector component. We conclu
de that vector analyses can provide a useful visualization of treatment eff
ects on tree growth and nutrition, Its relevance to the assessment of susta
inable forest management is limited by the need to have treatment compariso
ns available and interpretations should be corroborated by other evidence b
efore firm conclusions can be drawn. (C) 1999 Elsevier Science B.V. All rig
hts reserved.