Introducing the effect of extinction into the so-called replicator equation
s in mathematical biology, we construct a general model where the diversity
of species, i.e., the dimension of the equation, is a time-dependent varia
ble. The system shows very different behavior from the original replicator
equation, and leads to mass extinction when the system initially has high d
iversity. The present theory can serve as a mathematical foundation for the
paleontologic theory for mass extinction. This extinction dynamics is a pr
ototype of dynamical systems where the variable dimension is inevitable. [S
1063-651X(99)06707-0].