The spatial pattern of a small cattle herd within a fenced grazing pasture
was investigated using a mathematical model. Only changes in the distance b
etween the two most distant individuals in the pasture, i.e. troop length,
were observed. Troop length is a measure of closeness of individuals within
a herd. In the model, we assumed that the troop length was determined by:
(1) attractive activities (desire to be in a group) and repulsive activitie
s (maintenance of individual space) operating among individuals in the herd
; and (2) random activities unrelated to the attractive and repulsive activ
ities. A relationship between these two elements was incorporated into a di
ffusion equation, and the theoretical frequency distribution of the troop l
ength was derived. The model was tested using experimental data collected i
n a strip-wise pasture. Activities of a cattle herd were visually divided i
nto three categories such as resting, feeding and moving states. Parameters
representing the attractive and repulsive activities, and the equilibrium
of troop length were estimated using the model for the three states. The fr
equency distribution of troop length was compared between the three states
and with the frequency distribution derived under the assumption of a rando
m spatial pattern. It was concluded that: (1) the changes in troop length c
ould be described by the newly derived model; (2) the spatial pattern of a
cattle herd in an actual pasture was not random but aggregated; and (3) the
closeness of individuals within a herd was loosened in the following order
: resting, feeding and moving states. (C) 1999 Elsevier Science B.V. All ri
ghts reserved.