H. Utsugi, Angle distribution of foliage in individual Chamaecyparis obtuse canopies and effect of angle on diffuse light penetration, TREES, 14(1), 1999, pp. 1-9
The vertical distribution of foliage angle and area of three Chamaecyparis
obtusa trees was determined by the triangle method, which calculates foliag
e geometry using measured coordinates of the leaf "corners", in a 43-year-o
ld plantation in central Japan. Vertical distribution patterns of leaf area
were different depending on tree size, but the boundary heights, which div
ide the canopy into sunlit and shaded parts, were similar in the three samp
le trees. The value of the average foliage angle [I(Z)] at a given depth (Z
) from the tip of the stem decreased continually from the upper to lower la
yers within the canopy. The vertical patterns of changes in I(Z) were diffe
rent among the three trees, but could be expressed by the following allomet
ric equation as a function of depth.
I (Z)=aZ(b) exp(cZ)
where a, b and c are constants. The average foliage angle of C. obtusa depe
nded on the position within the canopy and tree size; the value was larger
in the sunlit parts of the canopy than in the shaded parts. However, the fo
liage angle distribution in the overall canopy fitted an ellipsoidal area d
istribution model. The probability of diffuse light penetration through the
canopy was calculated using foliage angle and cumulative leaf area paramet
ers. The probability was different from that calculated by Beer's Law for l
ight extinction, especially in the sunlit part of the canopy. These results
suggested that the foliage angle distribution within the canopy is an impo
rtant factor in: (1) the estimation of the absorption of diffuse radiation:
and (2) evaluation of the amount of absorbed direct radiation in the canop
y of this forest.