Enzyme data from 12 previously published studies of haploid bryophyte popul
ations were subjected to tests of selective neutrality and population equil
ibrium. Altogether 280 samples were tested using the Ewens-Watterson test,
the Ewens exact test, and Chakraborty's test. The Ewens-Watterson and the E
wens exact tests revealed neutrality in 84.6% of the cases, whereas the Cha
kraborty test revealed neutrality in 100% of the cases. Neutrality may thus
explain the relatively high amounts of genetic variation found in these or
ganisms. Possible hidden population substructuring and nonrandom field samp
ling of genotypes in many of the studies could make Chakraborty's test more
reliable than the two others.