How do the actions of individual genes contribute to the complex morphologi
es of animals and plants? How widespread are these genes taxonomically? How
many genes are involved in the morphological differences observed between
species, and can we identify them? To what extent can empirical data and th
eory be reconciled? We provide an overview of some recent attempts to answe
r these questions, answers that have taken us to the threshold of understan
ding the mechanistic basis and evolutionary factors that underlie morpholog
ical innovation.