Extinction of populations occurs naturally, but global extinction rates are
accelerating, making understanding extinction a high priority for conserva
tion. Extinction in experimental populations of brine shrimp (Artemia franc
iscana) was measured to assess hypothesized extinction processes. Greater i
nitial population size, greater maximum population size supported by the en
vironment, and Lower variation in environmental conditions reduced the Like
lihood of extinction, as hypothesized. However, initial population size was
less important, and maximum population size and environmental variation we
re more important than often hypothesized. Unexpectedly, deterministic osci
llations in population size due to inherent nonlinear dynamics and overcrow
ding were as important or more important than hypothesized processes.