The teeth of every primate, living and extinct, are covered by a hard, dura
ble layer of enamel. This is not unique: Almost all mammals have enamel-cov
ered teeth. In addition, all of the variations in enamel structure that occ
ur in primates are also found in other groups of mammals. Nevertheless, the
very complexity of enamel and the variation we see in it on the teeth of l
iving and fossil primates raise questions about its evolutionary significan
ce. Is the complex structure of primate enamel adaptive? What, if anything,
does enamel structure tell us about primate phylogeny? To answer these que
stions, we need to look more closely at the characteristics of prismatic en
amel in primates and at the distribution of those characteristics, both in
relation to our knowledge of primate dental function and feeding ecology an
d from a phylogenetic perspective.