New studies are documenting trophic cascades in theoretically unlikely syst
ems such as tropical forests and the open ocean. Together with increasing e
vidence of cascades, there is a deepening understanding of the conditions t
hat promote and inhibit the transmission of predatory effects. These condit
ions include the relative productivity of ecosystems, presence of refuges a
nd the potential for compensation. However, trophic cascades are also alter
ed by humans. Analyses of the extirpation of large animals reveal loss of c
ascades, and the potential of conservation to restore not only predator pop
ulations but also the ecosystem-level effects that ramify from their presen
ce.