Pliomerina Chugaeva, 1958, a characteristic element of the peri-Gondwanan E
okosovopeltis-Pliomerina Province, occurs in Llanvirn and Caradoc strata of
the Argentine Precordillera. Three protaspid instars are present in the on
togeny of the Early Caradoc Pliomerina peripata new species. A species poss
ibly representing Pliomerella is a rare element in Early Caradoc strata of
the Las Aguaditas Formation. Prosopiscus is represented by a Llanvirn speci
es with a first record of protaspid larvae for the enigmatic Family Prosopi
scidae. Early growth stages support closest relations of prosopiscids to Ph
acopina. The presence of Pliomerina and Prosopiscus, along with the cheirur
id Pateraspis, demonstrates a peri-Gondwanan influence on the Precordillera
n fauna by the Llanvirn, and supports the proximity of the Precordillera te
rrane to the South American margin by that time.